date: 2013-07-04T07:59:06Z pdf:unmappedUnicodeCharsPerPage: 3 pdf:PDFVersion: 1.4 pdf:docinfo:title: doi:10.1098/rspb.2009.0088 xmp:CreatorTool: Royal Society access_permission:modify_annotations: true access_permission:can_print_degraded: true dcterms:created: 2009-02-18T13:48:32Z Last-Modified: 2013-07-04T07:59:06Z dcterms:modified: 2013-07-04T07:59:06Z dc:format: application/pdf; version=1.4 title: doi:10.1098/rspb.2009.0088 xmpMM:DocumentID: uuid:c42b72ff-1dd1-11b2-0a00-748c400958e2 Last-Save-Date: 2013-07-04T07:59:06Z pdf:docinfo:creator_tool: Royal Society access_permission:fill_in_form: true pdf:docinfo:modified: 2013-07-04T07:59:06Z meta:save-date: 2013-07-04T07:59:06Z pdf:encrypted: false dc:title: doi:10.1098/rspb.2009.0088 modified: 2013-07-04T07:59:06Z Content-Type: application/pdf X-Parsed-By: org.apache.tika.parser.DefaultParser meta:creation-date: 2009-02-18T13:48:32Z created: 2009-02-18T13:48:32Z access_permission:extract_for_accessibility: true access_permission:assemble_document: true xmpTPg:NPages: 9 Creation-Date: 2009-02-18T13:48:32Z pdf:charsPerPage: 1552 access_permission:extract_content: true access_permission:can_print: true producer: Acrobat Distiller 5.0.5 (Windows) access_permission:can_modify: true pdf:docinfo:producer: Acrobat Distiller 5.0.5 (Windows) pdf:docinfo:created: 2009-02-18T13:48:32Z