date: 2014-03-27T21:22:25Z pdf:unmappedUnicodeCharsPerPage: 1 pdf:PDFVersion: 1.5 pdf:docinfo:title: ppat.1004081 1..13 xmp:CreatorTool: 3B2 Total Publishing System 7.51n/W access_permission:modify_annotations: true access_permission:can_print_degraded: true dcterms:created: 2014-03-25T00:50:17Z Last-Modified: 2014-03-27T21:22:25Z dcterms:modified: 2014-03-27T21:22:25Z dc:format: application/pdf; version=1.5 title: ppat.1004081 1..13 Last-Save-Date: 2014-03-27T21:22:25Z pdf:docinfo:creator_tool: 3B2 Total Publishing System 7.51n/W CrossMarkDomains[1]: www.plospathogens.org access_permission:fill_in_form: true pdf:docinfo:modified: 2014-03-27T21:22:25Z meta:save-date: 2014-03-27T21:22:25Z pdf:encrypted: false dc:title: ppat.1004081 1..13 modified: 2014-03-27T21:22:25Z pdf:docinfo:custom:CrossMarkDomains[1]: www.plospathogens.org Content-Type: application/pdf X-Parsed-By: org.apache.tika.parser.DefaultParser meta:creation-date: 2014-03-25T00:50:17Z pdf:docinfo:custom:CrossmarkMajorVersionDate: 2014-04-17 created: 2014-03-25T00:50:17Z access_permission:extract_for_accessibility: true access_permission:assemble_document: true xmpTPg:NPages: 13 Creation-Date: 2014-03-25T00:50:17Z pdf:charsPerPage: 5307 access_permission:extract_content: true access_permission:can_print: true pdf:docinfo:custom:doi: 10.1371/journal.ppat.1004081 pdf:docinfo:custom:CrossmarkDomainExclusive: false producer: Acrobat Distiller 9.0.0 (Windows); modified using iText 5.0.3 (c) 1T3XT BVBA; modified using iText® 5.1.3 ©2000-2011 1T3XT BVBA CrossmarkDomainExclusive: false access_permission:can_modify: true pdf:docinfo:producer: Acrobat Distiller 9.0.0 (Windows); modified using iText 5.0.3 (c) 1T3XT BVBA; modified using iText® 5.1.3 ©2000-2011 1T3XT BVBA pdf:docinfo:created: 2014-03-25T00:50:17Z CrossmarkMajorVersionDate: 2014-04-17 doi: 10.1371/journal.ppat.1004081