date: 2018-08-07T13:44:27Z pdf:docinfo:custom:AAPL:Keywords: pdf:PDFVersion: 1.4 pdf:docinfo:title: Modern wolves trace their origin to a late Pleistocene expansion from Beringia xmp:CreatorTool: Word access_permission:can_print_degraded: true subject: dc:format: application/pdf; version=1.4 pdf:docinfo:creator_tool: Word access_permission:fill_in_form: true pdf:encrypted: false dc:title: Modern wolves trace their origin to a late Pleistocene expansion from Beringia modified: 2018-08-07T13:44:27Z cp:subject: pdf:docinfo:subject: pdf:docinfo:creator: Liisa Loog meta:author: Liisa Loog meta:creation-date: 2018-07-17T14:31:28Z created: 2018-07-17T14:31:28Z access_permission:extract_for_accessibility: true Creation-Date: 2018-07-17T14:31:28Z Author: Liisa Loog producer: Mac OS X 10.12.6 Quartz PDFContext pdf:docinfo:producer: Mac OS X 10.12.6 Quartz PDFContext pdf:unmappedUnicodeCharsPerPage: 0 Keywords: access_permission:modify_annotations: true AAPL:Keywords: dc:creator: Liisa Loog dcterms:created: 2018-07-17T14:31:28Z Last-Modified: 2018-08-07T13:44:27Z dcterms:modified: 2018-08-07T13:44:27Z title: Modern wolves trace their origin to a late Pleistocene expansion from Beringia xmpMM:DocumentID: uuid:21bb4594-1dd2-11b2-0a00-0309271dd700 Last-Save-Date: 2018-08-07T13:44:27Z pdf:docinfo:keywords: pdf:docinfo:modified: 2018-08-07T13:44:27Z meta:save-date: 2018-08-07T13:44:27Z Content-Type: application/pdf X-Parsed-By: org.apache.tika.parser.DefaultParser creator: Liisa Loog dc:subject: access_permission:assemble_document: true xmpTPg:NPages: 29 pdf:charsPerPage: 2489 access_permission:extract_content: true access_permission:can_print: true meta:keyword: access_permission:can_modify: true pdf:docinfo:created: 2018-07-17T14:31:28Z