date: 2014-07-08T14:11:45Z pdf:unmappedUnicodeCharsPerPage: 0 pdf:PDFVersion: 1.5 xmp:CreatorTool: Arbortext Advanced Print Publisher 9.1.510/W Unicode access_permission:modify_annotations: true access_permission:can_print_degraded: true dcterms:created: 2014-06-13T03:10:20Z Last-Modified: 2014-07-08T14:11:45Z dcterms:modified: 2014-07-08T14:11:45Z dc:format: application/pdf; version=1.5 xmpMM:DocumentID: uuid:67c74737-3476-476b-b942-ab037b050b47 Last-Save-Date: 2014-07-08T14:11:45Z pdf:docinfo:creator_tool: Arbortext Advanced Print Publisher 9.1.510/W Unicode CrossMarkDomains[1]: www.pnas.org access_permission:fill_in_form: true pdf:docinfo:modified: 2014-07-08T14:11:45Z meta:save-date: 2014-07-08T14:11:45Z pdf:encrypted: false modified: 2014-07-08T14:11:45Z pdf:docinfo:custom:CrossMarkDomains[1]: www.pnas.org Content-Type: application/pdf X-Parsed-By: org.apache.tika.parser.DefaultParser meta:creation-date: 2014-06-13T03:10:20Z pdf:docinfo:custom:CrossmarkMajorVersionDate: 2014-06-13 created: 2014-06-13T03:10:20Z access_permission:extract_for_accessibility: true access_permission:assemble_document: true xmpTPg:NPages: 10 Creation-Date: 2014-06-13T03:10:20Z pdf:charsPerPage: 6061 access_permission:extract_content: true access_permission:can_print: true pdf:docinfo:custom:doi: 10.1073/pnas.1409567111 pdf:docinfo:custom:CrossmarkDomainExclusive: false producer: Acrobat Distiller 10.1.8 (Windows) CrossmarkDomainExclusive: false access_permission:can_modify: true pdf:docinfo:producer: Acrobat Distiller 10.1.8 (Windows) pdf:docinfo:created: 2014-06-13T03:10:20Z CrossmarkMajorVersionDate: 2014-06-13 doi: 10.1073/pnas.1409567111