date: 2016-01-11T07:33:47Z pdf:unmappedUnicodeCharsPerPage: 0 pdf:PDFVersion: 1.4 xmp:CreatorTool: Arbortext Advanced Print Publisher 9.1.510/W Unicode access_permission:modify_annotations: true access_permission:can_print_degraded: true dcterms:created: 2014-06-19T01:32:21Z Last-Modified: 2016-01-11T07:33:47Z dcterms:modified: 2016-01-11T07:33:47Z dc:format: application/pdf; version=1.4 xmpMM:DocumentID: uuid:1298aeef-a5a7-4f16-a212-ea9f59aec3bb Last-Save-Date: 2016-01-11T07:33:47Z pdf:docinfo:creator_tool: Arbortext Advanced Print Publisher 9.1.510/W Unicode CrossMarkDomains[1]: www.pnas.org access_permission:fill_in_form: true pdf:docinfo:modified: 2016-01-11T07:33:47Z meta:save-date: 2016-01-11T07:33:47Z pdf:encrypted: false modified: 2016-01-11T07:33:47Z pdf:docinfo:custom:CrossMarkDomains[1]: www.pnas.org Content-Type: application/pdf X-Parsed-By: org.apache.tika.parser.DefaultParser meta:creation-date: 2014-06-19T01:32:21Z pdf:docinfo:custom:CrossmarkMajorVersionDate: 2014-06-19 created: 2014-06-19T01:32:21Z access_permission:extract_for_accessibility: true access_permission:assemble_document: true xmpTPg:NPages: 6 Creation-Date: 2014-06-19T01:32:21Z pdf:charsPerPage: 6509 access_permission:extract_content: true access_permission:can_print: true pdf:docinfo:custom:doi: 10.1073/pnas.1403466111 pdf:docinfo:custom:CrossmarkDomainExclusive: false producer: Acrobat Distiller 10.1.8 (Windows) CrossmarkDomainExclusive: false access_permission:can_modify: true pdf:docinfo:producer: Acrobat Distiller 10.1.8 (Windows) pdf:docinfo:created: 2014-06-19T01:32:21Z CrossmarkMajorVersionDate: 2014-06-19 doi: 10.1073/pnas.1403466111